The recycling and exploitation of brewing residues, such as brewery wastewater, surplus yeast, and brewers' spent grain (BSG), are critical goals for reduction of energy consumption and residue disposal costs, and also for reduction of the associated carbon foot-print (Zupančič et al. 2017).BSG is the most abundant by-product from the brewing process (i.e., 85% of total by-products) (Fig. 1).